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2. STOP n° 2 : Triassic of Dio ; (GG, J Ga).
*
Stratigraphy (GG, fig. 6).
In the Lodévois area, Triassic series is about 250 m thick. Near Dio
village it overlays the Permian (Saxonian) with some degrees of unconformity.
Beneath the contact, in Permian red pelites, one can see numerous carbonate
nodules (dolocretes). These structures result from pedogenesis occurring before
the Triassic deposits.
According to ALABOUVETTE et al. (1982, 1987), the Triassic series consists
of the following formations from the base to the top (fig. 3).
– t3 :‘Grès de base’ (Lower
Sandstones = 30-50m) : alternation sandstones/red mudstones with sometimes pebbles-conglomerats
coming from the basement.
– t4-5 : ‘Argilites inférieures’ (Lower
Mudstones = 20-30m) with a succession of variegated mudstones (containing NaCl
pseudomorphose) / sandstones / dolostones yielding numerous footprint levels
(see below) and Voltzia type macroflora alike to the Lower Triassic macroflora
of the Vosges.
– t6 : ‘Grès médians’ and ‘Barre
carbonatée’ (Upper Sandstones and Conglomerates + Laminated Dolomite
= 80 m). Successively, this formation consists of dolomite / sandstone, massive
sandstones banks with conglomerates (50m) then dolomite.
Grès de base – Grès médians (Upper Sandstones and
Conglomerates) are the ‘Lower Formation’ sensu Lopez & Mader
(1992). This latter is dated Middle Anisian-Ladinian and contains footprints,
paleoflora and Myophoria sp. It indicates a playa-sabkha environment with occasional
fluvial channels.
– t7-9 : ‘Argiles Supérieures’ (Upper
Mudstones =100m). They are multicoloured with successively : carbonate interbedding,
Cavernous Dolomite, mudstones with anhydrite and gypsum.
The Formation t7-9 = former Keuper = Middle and Upper Formation sensu Lopez & Mader
(1992). It is dated Carnian (by palynoflora) and Norian.
– t10 : Rhétien = Rhaetian Formation. There
are 2 Members. The lower, 30 m thick, has sandstones, dolomites with fishes scales,
pelecypodes and Avicula contorta. The Upper Member (15 m) contains above all
carbonates.
In summary the Formations ‘Grès de base – Argilites inférieures’ (Lower
Sandstones-Lower Mudstones) form the lower megacycle meaning a vertical environmental
evolution from a fluvial system to a playa via a flood plain. The ‘Barre
Carbonatée’ and ‘Argiles Supérieures’ (Upper
Mudstones) Formations are the upper megacycle. It was deposited in an littoral
environment with supratidal sabkhas. Rhaetian is the last fluvial system before
the inter-subtidal marine hettangian sedimentation.

Fig 6– A
= Triassic serie after Alabouvette et al 1987 (CB : carbonated bank, F :
footprints). F = traces de pas) ; B à I = ichnites avec B et D = piste
Rotodactylus bessieri DEMATHIEU, 1984, C = Squelette pédieux de Macrocnemus
bassanii, E = Brachychirotherium gallicum (WILLRUTH, 1917) : 2 couples pied
(P)-main (M), F = squelette pédieux reconstitué, G = pistes
partielles de Brachychirotherium gallicum, H = Chirotherium barthii, I =
piste Rhynchosauroides lutevensis DEMATHIEU, 1984 ;
* Palaeontology
• Palichnofauna (GD, fig.
6)
Vertebrate fossils have not been discovered into these Triassic layers. In
contrast numerous ichnofossils have been mentioned and some of them, believed
to be derived from vertebrates, have been known since 1857 and first described
by P. GERVAIS. They include casts on the lower surface of thick slabs that
come from the neighbourhood of Lodève, the village of Fozières.
No one of these slabs, supplied to different Museums, Paris, Lodève,
Genève, Lyon etc., has been founded in situ. Nevertheless it seems probable
that their stratigraphic position lies into ‘t.4-5’. These levels
contain the following footprints : Chirotherium barthii Kaup
1835, Brachychirotherium gallicum (WILLRUTH, 1917), Brachychirotherium
circaparvum DEMATHIEU, 1971, Rotodactylus bessieri DEMATHIEU,
1984, Rhynchosauroides peabodyi FABER 1958, Rhynchosauroides
lutevensis DEMATHIEU, 1984.
1 – Chirotherium barthii (221 x 155 mm) is represented by only a few footprints and was probably derived from a quadrupedal and pentadactyl Thecodont whose foot structure was somewhat advanced in the evolution of the Archosaurs by the high value of the cross axis (=90°), a very narrow trackway, and a high value of the pace angulation (about 180°).
2 – Brachychirotherium
gallicum (WILLRUTH, 1917) (B.g, fig. 6A) is smaller
(175 x 110mm) and the pes shows other characters : cross-axis is ≤ 70° and
so indicates a more primitive organization of the basipode of the foot.
Cross-axis lies between 70-80° – a common value for the ichnospecies
of this genus. The fifth toe is short and directed straight to the
outer part of the pes. Small and round scales are visible on a very
well preserved imprint. A manus is always present and is smaller than
the pes, by a quarter. It has about the same shape but its width is
greater than the length.
3 – Brachychirotherium
circaparvum DEMATHIEU,
1971 (B. C
fig. 6A). Like
the other pes
imprints of
this genus,
the digits
are rather
thick with
claws directed
toward the
outer side
of the pes,
except the
fourth for
which it seems
it has no claw.
Among digits,
II ≤ III.
The manus is
somewhat rotated
toward the
outer side
of the trackway.
The characters
of these footprints
lead us to
think they
were made by
a thecodont
sphenosuchien.
4 – Synaptichnium
sp.
NOPCSA, 1923.
This ichnogenus
is known here
from only two
specimens ;
its main characters
are : quadrupedy,
pentadactyly,
subparallelism
of the four
first toes.
The IVth is
the longest.
The fifth is
right and in
the continuity
of the metatarsal.
The cross-axis
is low : 60-70° which
points out
a primitive
feature of
the basipode
.
5 – Rotodactylus is
a very peculiar
pentadactyl
imprint showing
the fifth.
Digit V impressed
behind the
set of the
four first
digits. Rotodactylus
bessieri DEMATHIEU,
1984 has a
medium size
for the ichnogenus.
The trackmaker
is not well
known ; Dinosaur
or Lepidosaur
according to
different authors.
6 – Rhynchosauroides
lutevensis DEMATHIEU,
1984 is a very
frequent species
in the lower
and middle
Triassic. The
imprint of
the pentadactyl
foot shows
only three
toes, the II,
III ,IV, the
others being
too short to
mark the soil
.The pentadactyl
manus lies
behind the
pes and shows
four sub-parallel
straight digits,
the fifth is
located behind
them. The trackmaker
is of small
size (60-70cm),
and was probably
a Lepidosaur.
The manus lies
behind the
pes and indicates
long limbs.
These footprints
appear in the
red-violet
horizon.
7– Rhynchosauroides
peabodyi, FABER
1958.
This ichnospecies is large and presents some peculierities : manus and pes
have the same shape ; digits are well separated, long, curved with accurate
claws. Like the other ichnospecies the manus is put behind the pes.
Because the homopody and the presence of round scales under the toes, it is
probable that the trackmaker was an archosaur like Chasmatosaurus
8– Rhynchosauroides
cf. petri DEMATHIEU,
1966. This
small ichnospecies
is very common
in the Triassic
of the eastern
border of the
Massif Central.
It is very
small and its
characters
are the same
as those of
other ichnospecies.
It is possible
that these
footprints,
which have
the same shape,
are the result
of animals
of the same
species at
different stages
of growth ?
• Paleoflora (JGa, JB)
DOUBINGER & CHEYLAN (1964) described a small plant assemblage with Voltzia heterophylla, Albertia elliptica, Equisetites mougeottii, Schizoneura cf. paradoxa, Endolepis vogesiaca and Aethophyllum sp. with a microflora characterized by Illinites kosankei, Chordasporites singulichorda and Triadispora falcata suggestive of a lower Anisian age. A more rich microflora allowed DOUBINGER & ADLOFF (1981) to propose a middle Anisian age ; this was confirmed by the recent study of DIEZ FERRER (2000) in the area north of La Lieude where a rich microflora with the association Illinites kosankei, I. chitonoides, Concentricisporites nevesi, Microcachryidites sittleri indicates an early Pelsoniense age.
Between
stops 2 and 3 after the Merquière pass, a general
view allows us to see the Triassic series resting on the Permian red
pelites dipping at about 15° to the East.